About 3.6 million years ago

Walking upright

Footprints pressed into volcanic ash in Tanzania show a human-like stride, long before our own genus appeared.

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What the evidence shows

  • The Laetoli footprints in Tanzania, about 3.6 million years old, are the earliest direct evidence of hominins walking on two legs.
  • Experiments comparing them with modern footprints suggest their makers walked with extended, human-like legs rather than the bent-knee gait of apes.

What is uncertain

  • Why walking upright evolved is not settled. Saving energy was probably one important pressure.
  • The prints are usually attributed to Australopithecus afarensis, but footprints do not name their maker.

Sources

Summaries here are our own. Some publishers limit access to full papers; learning on this site is never paywalled.